riers and environmental instabilities inherent to marginal bays and seaways). The two marine bioprovinces differed dramatically in type of dominant shellfish. This is consistent with recent paleogeographic reconstructions that suggest a lack of a direct marine connection between the Paratethys and Boreal Province in the Middle Miocene. The study shows also that the value of the biological information entombed in the fossil record can be improved by employing an hierarchical approach, in which differences among samples of fossils are evaluated quantitatively as a function of the geographic and temporal scales of observation.

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Contact: Ann Cairns
acairns@geosociety.org
303-357-1056
Geological Society of America
8-Feb-2002

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